Denisovans were an archaic hominin species that was a sister to Neanderthals and admixed with modern humans in Southeast Asia. Previously, Denisovan remains were found in Siberia and Tibet.
Now, the tooth of a Denisovan female which is at least 131,000 years old, was found in Cobra Cave in Laos. The species identification and gender were based upon morphology and tooth enamel proteins, rather than ancient DNA per se. The presence of Denisovan's in the region was expected based upon Denisovan DNA in Asians and Australians and Papuans. The New York Times discusses the find further.
The paper and its abstract are as follows:
The Pleistocene presence of the genus Homo in continental Southeast Asia is primarily evidenced by a sparse stone tool record and rare human remains. Here we report a Middle Pleistocene hominin specimen from Laos, with the discovery of a molar from the Tam Ngu Hao 2 (Cobra Cave) limestone cave in the Annamite Mountains. The age of the fossil-bearing breccia ranges between 164–131 kyr, based on the Bayesian modelling of luminescence dating of the sedimentary matrix from which it was recovered, U-series dating of an overlying flowstone, and U-series–ESR dating of associated faunal teeth. Analyses of the internal structure of the molar in tandem with palaeoproteomic analyses of the enamel indicate that the tooth derives from a young, likely female, Homo individual. The close morphological affinities with the Xiahe specimen from China indicate that they belong to the same taxon and that Tam Ngu Hao 2 most likely represents a Denisovan.
Fabrice Demeter, et al.,"A Middle Pleistocene Denisovan molar from the Annamite Chain of northern Laos" 13 Nature Communications volume 13, Article number: 2557 (May 17, 2022) (open access).